| 20 August 1910 | Born London, England |
| 1931 | Educated University of London BSc (Hons) |
| 1932 | University of London, MSc |
| 1932 - 1935 | Assistant Lecturer in Zoology, University College London |
| 1935 - 1940 | Lecturer in Zoology |
| 1937 - 1938 | Rockefeller Research Fellow, Columbia University, United States of America |
| 1940 | University of London, DSc |
| 1940 - 1945 | Wartime positions as Statistician and Entomologist, British Ministry of Food |
| 1945 - 1947 | Reader in Zoology, University of London |
| 1947 | Guest Investigator, Department of Genetics, Carnegie Institute of Washington, Cold Spring Harbor, New York |
| 1947 - 1953 | Professor of Zoology, University of Texas |
| 1953 - 1956 | Senior Research Fellow, Genetics Section, Division of Plant Industry, Commonwealth Scientific and Industrial Research Organization, Canberra |
| 1955 | Fellow of the Australian Academy of Science |
| 1957 - 1958 | Associate Professor, then Professor of Zoology, University of Missouri |
| 1958 - 1964 | Professor of Zoology, University of Melbourne |
| 1959 | MSc, University of Melbourne |
| 1961 | Fellow of the Royal Society |
| 1962 | President, Section D, Australian and New Zealand Association for the Advancement of Science |
| 1963 | Honorary Foreign Member, American Academy of Arts and Sciences |
| 1964 - 1975 | Professor of Genetics, University of Melbourne |
| 1965 | Mueller Medal, Australian and New Zealand Association for the Advancement of Science 1965 |
| 1969 - 1971 | President, Australian Entomological Society |
| 1970 - 1972 | President, Genetics Society of Australia |
| 1976 - 1983 | Visiting Fellow, Department of Population Biology, Australian National University |
| 1978 | Foreign Member, American Philosophical Society |
| 1978 | Socio Straniero, Accademia Nazionale dei Lencei |
| 1979 | Silver Medal for Research, Royal Society of Victoria |
| 1980 | Honorary Member |
| 1981 | Foreign Associate, United States National Academy of Sciences |
| 1983 | Minerva Medal, University of Rome |
| 1983 | Linnean Medal for Zoology, Linnean Society of London |
| 16 December 1983 | Died Canberra |