| 5 March 1938 | Born United Kingdom |
| 1959 | BSc (Hons), University of Edinburgh |
| 1962 | PhD, University of Birmingham |
| 1963 - 1964 | NATO Research fellowship, CSIRO Division of Plant Industry |
| 1964 - 1970 | Lecturer in botany, Department of Botany, School of General Studies, Australian National University |
| 1974 - 1976 | Chairman, Victorian branch, Australian Society of Plant Physiologists |
| 1974 - 1997 | Senior lecturer 1970-73, Professor of Botany, University of Melbourne |
| 1976 | President, Botany section, ANZAAS Hobart congress |
| 1977 | Chairman, Botany section, ANZAAS, Melbourne |
| 1982 - 1987 | Co-director, Plant Cell Biology Research Centre, University of Melbourne |
| 1984 | DSc, University of Melbourne |
| 1988 - ? | Deputy chairman, School of Botany |
| 1989 | Fellow of the Australian Academy of Science |
| 1990 - 1997 | President, International Association for Sexual Plant Reproduction |
| 1995 | Co-editor of Biology, a textbook aimed at first year undergraduates |
| 30 August 1997 | Died Melbourne |